Interestingly almost all of these subtle

The extent of morphological and behavioural variation within colonies was also associated with their speed of emergence and attack, but only in field conditions. Colonies composed of a more behaviourally diverse set of individuals, in terms of their boldness, were faster to emerge in response to prey. Whereas, colonies composed of more morphologically diverse individuals, in terms of their prosoma width, were faster to attack. Although, again, the reasons for these effects are unknown, we propose that Bikinin trait complementarity and catalytic interactions between unlike individuals may cause changes in colonies' collective behaviour. For instance, across many animal societies, individuals may change their behaviour based on the behavioural tendencies (Harcourt et?al., 2009, Holbrook et?al., 2014 and King et?al., 2015) or the body sizes (Amir et al., 2000) of their fellow group members which, in turn, could impact the collective behaviour of the group (Jandt et?al., 2014 and Modlmeier et?al., 2014b). Finally, as seen in other studies (Dyer et?al., 2009, Hui and Pinter-Wollman, 2014 and Keiser and Pruitt, 2014), creating colonies of contrasting personality compositions had impacts on collective behaviour. In bilateral symmetry study, colonies composed of bold individuals responded to prey with twice as many attackers.